Riaselis rugosa E.M.Friis, P.R.Crane et K.R.Pedersen

Plant Fossil Names Registry Number: PFN000135

Act LSID: urn:lsid:plantfossilnames.org:act:135

Authors: E. M. Friis, P. R. Crane & K. R. Pedersen

Rank: species

Genus: Riaselis E.M.Friis, P.R.Crane et K.R.Pedersen

Reference: Friis, E. M., Crane, P. R. & Pedersen, K. R. (2019): Extinct Diversity among Early Cretaceous Angiosperms: Mesofossil Evidence of Early Magnoliales from Portugal. – International Journal of Plant Sciences 180(2): 93–127.

Page of description: 113

Illustrations or figures: figs 21, 22

Name is type for

Riaselis E.M.Friis, P.R.Crane et K.R.Pedersen 2019


Holotype S174029 (Famalicão 25), Palaeobotanical Collections, Department of Palaeobiology, the Swedish Museum of Natural History, Stockholm, Sweden
Figures: figs 21A–E, 22A–C

Note: Paratypes: S107711–S107717, S170241, S170242, S174030, S174422, S174825, S175196–S175198 (Famalicão sample 25).

Original diagnosis/description

Fruit/fruitlet indehiscent, with a thin wall; one seeded. Style lacking. Inner layer of fruit wall with narrow, elongate fibers. Stigmatic area restricted. Seed small, anatropous, bitegmic, and mesotestal-endotestal, with thick outer integument (testa) and thin inner integument (tegmen). Mesotesta one cell layer thick, composed of larger sclerenchyma cells. Endotesta several cell layers thick, composed of small polygonal cells with endoreticulate infillings. Seed surface coarsely rugulate. Micropyle formed from inner integument. Hilum and micropyle close together, not separated by testal tissue, resulting in a wide transversely oriented micropylar-hilar opening in the testa. Ovular vascular bundle continuing beyond the chalaza on the antiraphal side of the seed to the micropyle. Embryo tiny. Endosperm cellular, not-ruminate.


From the rugose surface of the seed (Latin rugosus [wrinkled])


Cretaceous, Lower Cretaceous
below the Figueira da Foz Formation; Early Cretaceous (late Aptian–early Albian or older)


Famalicão (lat. 39°42′16″N, long. 8°46′12″W)

Plant fossil remain

macro- and meso-fossils-embryophytes except wood


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