Dicotylophyllum expansolobum Upchurch et Dilcher

Plant Fossil Names Registry Number: PFN001493

Act LSID: urn:lsid:plantfossilnames.org:act:1493

Authors: G. R. Upchurch & D. L. Dilcher

Rank: species

Reference: Upchurch, G. R., Jr. & Dilcher, D. L. (1990): Cenomanian angiosperm leaf megafossils from the Rose Creek locality of the Dakota Formation, southeastern Nebraska. – Bulletin of the United States Geological Survey 1915: 1–55, 31 pls., link

Page of description: 48

Illustrations or figures: text-fig. 25, pl. 31


Holotype UF15713-8304, 8304', Paleobotany Collections, Florida Museum of Natural History, Gainesville, Florida, USA
Figures: text-fig. 25, pl. 31

Note: Holotype is formed by part and counterpart.

Original diagnosis/description

Leaf simple, five-lobed; lobes all diverging at approximately the same level on the leaf; each lobe widest above the base; margin with minute teeth that are difficult to see without magnification. Primary venation basally palmate; primary veins 5, stout/massive.

Emended diagnosis

Wang and Dilcher (2018: Palaeont. Electr., 21(3): 34):
Leaf simple, irregularly fivelobed, margin toothed near apex of lobes, serrations minute, simple. Petiole long and thin, enlarged both distally and proximally. Primary venation basal actinodromous. Secondary venation predominately brochidodromous or semicraspedodromous when margin is toothed near apex; typically, one series of loops present in the excostal region if margin is entire; secondary veins below sinus and between adjacent primary veins forming inverted ‘V’ pattern. Intersecondary veins common, composite. Tertiary and quaternary veins orthogonal reticulate, forming predominately quadrangular meshes.


Latin, expansus = spread out, and lobus =lobe, referring to the expansion of the lobes above their base.


Cretaceous, Upper Cretaceous, Cenomanian
Dakota Formation


United States
Rose Creek I locality, Nebraska [locality name according to Wang and Dilcher (2018: Palaeont. Electr., 21(3): 35]

Plant fossil remain

macro- and meso-fossils-embryophytes except wood


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