Tumidopteris astra Naugolnykh

Plant Fossil Names Registry Number: PFN001847

Act LSID: urn:lsid:plantfossilnames.org:act:1847

Author: S. V. Naugolnykh

Rank: species

Genus: Tumidopteris Naugolnykh

Reference: Naugolnykh, S. V. (2020): A new species of the genus Tumidopteris Naugolnykh from the Permian of the Pechora Cis-Urals, Russia. – Fossil Imprint 76(2): 270–278.

Page of description: 271

Illustrations or figures: text-figs 2a–e, 3a–e, 4a–f, 5a, b


Holotype GIN 4851/343h, Monographic Department of the State Darwin Museum, Moscow, Russian
Figures: text-figs 2a, d, 3a–e, 4a–c, e, 5a–c

Note: During study, the collection is kept in the Geological Institute of Russian Academy of Sciences (Moscow, Russia).

Original diagnosis/description

Fronds at least bipinnate. Fertile pinnules from pecopteroid to sphenopteroid, of subtriangular shape, with acute apices. Venation pinnate, with thin undulating midvein and three to five pairs of lateral veins. Lateral veins simple to dichotomizing up to three times. Terminations of lateral veins on fertile pinnules bear rosette-like sori. Each sorus consists of six to seven sporangia fused by their basal parts. Sporangia of short-clavate or prolonged ovoid shape, with small shallow sinus at apex, with wide indistinct annulus and vertical split for releasing spores. Sterile pinnules similar to fertile ones, but lacking sporangia. Associated spores round to ovoid, with small trilete mark.
Measurements of morphological structures are as followed: largest pinnule size: 7 mm long and 4 mm wide; average size of sporangia: 500 μm long, 300 μm wide; size of sporangial cells: 150 μm long, 60–70 μm wide; number of rows of cells in the annulus: presumably 1.


From “astra” (Latin for “star”; as in: “Per aspera ad astra”); after the star-like shape of the sori in this new species.


Permian, Guadalupian, Roadian


Russian Federation
Pechora coalbasin, the borehole IK-675, depth 961.7 m

Plant fossil remain

macro- and meso-fossils-embryophytes except wood


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