Cardstonia tolmanii M.Riley et Stockey

Plant Fossil Names Registry Number: PFN001880


Authors: M. G. Riley & R. A. Stockey

Rank: species

Genus: Cardstonia M.Riley et Stockey

Reference: Riley, M. G. & Stockey, R. A. (2004): Cardstonia tolmanii gen. et sp. nov. (Limnocharitaceae) from the Upper Cretaceous of Alberta, Canada. – International Journal of Plant Sciences 165(5): 897–916., link

Page of description: 899

Illustrations or figures: figs 2–3

Name is type for

Cardstonia M.Riley et Stockey 2004


Holotype UAPC-ALTA S55138, University of Alberta (UAPC-ALTA), Edmonton, Canada
Figures: Fig. 2a

Note: Paratypes: UAPC-ALTA S50947, S52263, S52279, S52268, S52266, S52272, S50989.

Original diagnosis/description

Leaves simple, ovate to elliptic, 5–12 cm long, 3.5–8.5 cm wide, l:w ratio 1.4:1, margin entire; apex convex to rounded; base cordate. Petiole 3–4 mm wide, at least 2 cm long, petiolar attachment marginal. Venation campylodromous, primary veins 23–27, with 3–5 medial veins originating at or near base; outer primaries running in strongly recurved arches that converge apically, merging at or near apex; apical pore present. Prominent fimbrial vein present. Secondary veins diverging at angles 45°–60° near midvein and 90° near margin; alternating major and minor secondary veins, with variable ABAB pattern, occasionally anastomose and dichotomize. Tertiary (transverse) veins few, with straight to slightly curved courses, forming moderately developed (irregular shape, variable-sized) areolae.


For Shayne Tolman of Cardston, Alberta, who brought the locality to the attention of the authors and spent many hours working with them in the field.


Cretaceous, Upper Cretaceous
Late Campanian–Early Maastrichtian
St. Mary River Formation


Along the banks of the St. Mary River below the St. Mary Reservoir dam and Spillway near Cardston, Alberta (T5 R254 S12 SE 1/4 W4M; 49°22′10″N, 113°06′10″W; UTM Grid 12: 5470600N, 347300E)

Plant fossil remain

macro- and meso-fossils-embryophytes except wood


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