Todites emarginatus Naugolnykh

Plant Fossil Names Registry Number: PFN001920


Author: S. V. Naugolnykh

Rank: species


Reference: Naugolnykh, S. V. (2013): Permian Ferns of Western Angaraland. – Paleontological Journal 47(12): 1379–1462., link

Page of description: 1408

Illustrations or figures: pl. 14, figs 1–3, pl. 15, fig. 5


Holotype 4851/173, Geological Institute, Russian Academy of Sciences, Moscow, Russia
Figures: figs 10j, l, 11, 12, pl. 14, figs 1–3, pl. 15, fig. 5

Original diagnosis/description

Fronds at least bipinnate. Sterile pinnules ranging from pecopteroid to sphenopteroid, with lobate margins and round apices. Pinnule midveins slightly undulating. Lateral veins dichotomizing up to two times. Fertile pinnules possessing isolated free sporangia, sometimes arranged in compact clusters. Thick-walled cells specialized for opening stomium located above stomium and forming short row. Each fertile pinnule with 80–100 sporangia.


From the Latin emarginatus (emarginate, sinuate), based on the dissected leaf lamina of pinnules into round lobes.


Upper Permian, Severodvinian Stage


Russian Federation
Pechora Coal Basin, Pechora River, outcrop no. 22 (after Zalessky, 1934a)

Plant fossil remain

macro- and meso-fossils-embryophytes except wood


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