Todites coronatus Naugolnykh

Plant Fossil Names Registry Number: PFN001921

Act LSID: urn:lsid:plantfossilnames.org:act:1921

Author: S. V. Naugolnykh

Rank: species

Genus:

Reference: Naugolnykh, S. V. (2013): Permian Ferns of Western Angaraland. – Paleontological Journal 47(12): 1379–1462., link

Page of description: 1412

Illustrations or figures: pl. 16, figs 1–10, pl. 17, figs 1–6

Types

Original diagnosis/description

Fern with fertile or partly fertile last order pinnae with pecopteroid pinnules. Frond rachis thick, robust, having central adaxial furrow. Sterile pinnules pecopteroid, connected by their bases and forming limb (wing), surrounding rachis of last order pinna. Pinnule midveins distinct, extending for three-fourths of pinnule extent and then dichotomizing once or twice (in basal part of pinna), only once (in middle part of pinna), or simple (in apical part of pinna). Pinnule apex round or blunted. Fertile pinnules pecopteroid, but sometimes constricted at base. Each fertile pinnule with up to 50 sporangia positioned on pinnule abaxial surface. Sporangia clavate or fusiform, consisting of two cell types. Basal and middle parts of sporangium consisting of relatively small elongated cells with thin walls. Apical part of sporangium consisting of larger cells with thick walls. Thick-walled apical cells forming crownlike structure surrounding stomium of sporangium and responsible for opening stomium. Spores in situ of Osmundacidites type, ranging
from round to subtriangular, up to 50 μm in diameter, with exine having small spines.

Etymology

From the Latin coronatus (crowned), based on the presence of well-developed specialized cells with thickened walls on the sporangium apex.

Stratigraphy

Permian
Middle Permian, Kazanian Stage

Locality

Russian Federation
Pechora Coal Basin, borehole IK 657, 532.0 m of depth