Cryptocaryoxylon grandoleaceum Akkemik

Plant Fossil Names Registry Number: PFN002211


Author: Ü. Akkemik

Rank: species

Reference: Akkemik, Ü. (2021): A re-examination of the angiosperm wood record from the early and middle Miocene of Turkey, and new species descriptions. – Acta Palaeobotanica 61(1): 42–94.

Page of description: 59

Illustrations or figures: pls 7, 8


Holotype GOK05, Department of Forest Botany, Faculty of Forestry, Istanbul University-Cerrahpasa, Istanbul, Turkey
Figures: pls 7, 8

Note: Holotype: Three thin sections of specimen GOK05.

Original diagnosis/description

This wood was identified as the fossil genus Laurinoxylon by Güngör et al. (2019), and is re-studied here. Growth ring boundaries distinct. Vessel diffuse to semiring-porous. Fibres thick-walled at growth ring boundary. Vessels arranged in no specific pattern, solitary, and radial multiples of 2–3 vessels. Mean tangential diameter of vessel lumina 50–100 μm, 20–40 vessels per mm2, vessel frequency decreasing in latewood (Pl. 7A, B). Axial parenchyma scanty paratracheal, diffuse, marginal (Pl. 7C, D), strand length 1–4 cells. Ray width 1–3 cells (Pl. 7E, F), 4–12 rays per mm. In tangential section, width of oil and mucilage cell same as ray width or more (Pl. 7E, F). Oil and mucilage cells associated with rays and among fibres (Pl. 7E, F). Intervessel pits alternate and small (4–7 μm). Perforation plates simple. Shape of alternate pits polygonal or circular (Pl. 8A). All rays procumbent with very large oil and mucilage cells, and body ray cells procumbent with 1 row (rarely 2 rows) of upright and/or square marginal cells and with very large oil and mucilage cells. Height of oil and mucilage cells 70–100 μm (Pls 7E, F, 9B–D). Axial parenchyma common in the wood (Pl. 8E).

PFNR Administrator note: mm2 to be read as square mm (our text box does not allow upper index).


The epithet “grandoleaceum” refers to the characteristics of the wood: very large oil and mucilage cells associated with rays and fibres.


Neogene, Miocene
Kesmekaya Volcanics, middle Miocene


East of Eşelek village – Gökçeada, near coastline

Plant fossil remain

fossil wood


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