Fraxinoxylon beypazariense Akkemik

Plant Fossil Names Registry Number: PFN002212


Author: Ü. Akkemik

Rank: species

Reference: Akkemik, Ü. (2021): A re-examination of the angiosperm wood record from the early and middle Miocene of Turkey, and new species descriptions. – Acta Palaeobotanica 61(1): 42–94.

Page of description: 64

Illustrations or figures: pls 10, 11


Holotype AGU13, Department of Forest Botany, Faculty of Forestry, Istanbul University-Cerrahpasa, Istanbul, Turkey
Figures: pls 10, 11

Note: Holotype: Three thin sections of specimen AGU13.

Original diagnosis/description

Growth ring boundaries distinct, wood ring-porous. Earlywood vessels wide, tyloses common. Vessels in latewood narrower. Vessels solitary and in radial multiples of up to three vessels in earlywood, vessel clusters common in latewood (Pl. 10A, B). Earlywood vessels sometimes in dendritic pattern. Tangential and radial diameter of vessel lumina 69 (26–116) μm and 90 (33–130) μm in earlywood, 27 (13–46) 50 μm and 24 (11–52) μm in latewood. Vessel density 83 (67–104) per mm2, with 26 (23–32) vessels per mm2 in earlywood and 57 (44–72) vessels per mm2 in latewood (Acarca Bayam et al., 2018). Paratracheal axial parenchyma almost lozenge-aliform, especially in latewood. 1–4 rows of marginal banded axial parenchyma present (Pl. 10A, B). Rays 1–2 (–3)-seriate, ≥12 per mm. Ray height 8–25 (–52) cells (Pls 10C, 11A). Intervessel pits alternate, small (4–7 μm), sometimes scalariform (Pl. 10D, E). Axial parenchyma in longitudinal section common, fibres thinto thick-walled (Pl. 11B). Perforation plates simple (Pl. 11C). Rays mainly heterocellular, body ray cells procumbent with 1–4 rows of upright and square marginal cells, sometimes homocellular with only procumbent body ray cells (Pl. 11D).

PFNR Administrator note: mm2 to be read as square mm (our text box does not allow upper index).


The epithet “beypazariense” refers to the origin of the sample. Beypazarı city has many fossil sites. This fossil genus has not previously been reported from Turkey.


Neogene, Miocene
Hançili Formation, early Miocene


Near Aşağıgüney village , Beypazarı, Ankara

Plant fossil remain

fossil wood


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