Carpites cordiformis Lesq.

Plant Fossil Names Registry Number: PFN000439


Author: C. L. Lesquereux

Rank: species

Reference: Lesquereux, C. L. (1892): The flora of the Dakota Group: A posthumous work. – Monographs of the United States Geological Survey 17: 1–400, 66 pls.

Page of description: 220

Illustrations or figures: pl. 22, fig. 9

Name is type for

Dakotanthus Manchester, Dilcher, Judd, Corder et Basinger 2018

Other combinations

Dakotanthus cordiformis (Lesq.) Manchester, Dilcher, Judd, Corder et Basinger 2018


Holotype USNM 50016, National Museum of Natural History, Smithsonian Institution, Washington, D.C., United States
Figures: pl. 22, fig. 9

Note: Holotype re-figured by Manchester et al. (2018: Acta Palaeobot., 58(1): pl. 3, fig. 11).

Holotype inventory number and repository according to Manchester et al. (2018: Acta Palaeobot., 58(1): 31).

Original diagnosis/description

Fruit deeply impressed into the stone, cordiform, separated in the middle by a deep, linear furrow, as though composed of two narrowly obovate ovules, which are straight and confluent in the middle, rounded above, curved on the sides, pointed at the upper end, convex on the surface.

Emended diagnosis

Manchester et al. (2018): Acta Palaeobot., 58(1): 32–33:
Flowers 22–30 mm diameter, 14–19 mm high, with two five-merous, hypogynous perianth whorls. Calyx thick, persistent; calyx lobes ovate, 10–13 mm long (from center of flower to apex of calyx lobe), 4.7–8.7 mm wide. Petals spathulate, membranous, 12–15 mm long, 10–15 mm wide, attached near base of gynoecium. Ovary dome-shaped, 8–10 mm wide, pentacarpellate with five locules. Five styles diverging from common point at apex of ovary, each with capitate stigma. Five crescent-shaped nectariferous pads situated at base of gynoecium and opposite and adnate to each sepal. Ten stamens inserted at level of the nectaries, one whorl organized opposite sepals (each at apex of a nectary lobe) and the other whorl opposite petals (and in sinus between adjacent nectary lobes). Stamens consisting of filament 5–7 mm long with flared base and dorsifixed ovate-ellipsoidal anther 3–6 mm long, 2.7–3.5 mm wide. In situ pollen oblate, triaperturate, grains oblate to spheroidal, 8–12 μm in equatorial diameter; apertures appear porate at surface but frequently somewhat torn meridionally, possibly along short colpi; ornamentation finely verrucate. Fruits globose to ovoid, five-valved loculicidal capsules, 10–12 mm wide and high, borne on thick, short pedicels (2 mm wide, 3 mm long) and with persistent sepals and persistent nectaries; capsules with thin septa, opening without central column.


Cretaceous, Lower Cretaceous, Albian
Dakota Formation (sandstone) [stratigraphy according to Manchester et al. (2018): Acta Palaeobot., 58(1): tab. 1]


United States
39°16.212′ N, 97°38.377′ W, Ottawa Co., Kansas [locality according to Manchester et al. (2018): Acta Palaeobot., 58(1): tab. 1]

Plant fossil remain

macro- and meso-fossils-embryophytes except wood


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