Lusitanispermum choffatii E.M.Friis, P.R.Crane et K.R.Pedersen

Plant Fossil Names Registry Number: PFN000097


Authors: E. M. Friis, P. R. Crane & K. R. Pedersen

Rank: species

Genus: Lusitanispermum E.M.Friis, P.R.Crane et K.R.Pedersen

Reference: Friis, E. M., Crane, P. R. & Pedersen, K. R. (2018): Extinct taxa of exotestal seeds close to Austrobaileyales and Nymphaeales from the Early Cretaceous of Portugal. – Fossil Imprint 74(1–2): 135–158.

Page of description: 147

Illustrations or figures: text-figs 7–10

Name is type for

Lusitanispermum E.M.Friis, P.R.Crane et K.R.Pedersen 2018


Holotype S174345 (Famalicão sample 025), Palaeobotanical Collections, Department of Palaeobiology, the Swedish Museum of Natural History, Stockholm, Sweden
Figures: text-figs 7a, b, 9a, b

Note: Paratypes: S105097, S105098, S170239, S174035, S174353, S174467-S174470, S174472, S174474, S174811, S175113-S175118 (Famalicão sample 025); deposited in Palaeobotanical Collections, Department of Palaeobiology, the Swedish Museum of Natural History, Stockholm, Sweden.

Original diagnosis/description

Isolated seeds occurring singly. Seeds small, anatropous, bitegmic and exotestal. Seeds bilaterally symmetrical with dorsiventral plane of symmetry. Seed surface smooth without longitudinal ridges. Raphe indistinct externally. Hilum and micropyle close together separated by a narrow zone of testal sclerenchyma. Hilar scar small without a hilar rim. Micropyle formed by the inner integument (tegmen) and marked on the seed surface by a transverse slit through the outer integument (testa) adjacent to the hilar scar. Testa formed from an outer layer (exotesta) of palisadeshaped sclerenchyma cells, and an inner thick layer of large parenchyma cells (mesotesta/endotesta). Palisade-shaped cells of exotesta with unevenly thickened walls that are thick towards the outside, thin towards the inside, and have a conical lumen. Exotesta thick adjacent to the hilum forming a bulging zone that may be two-parted. Anticlinal walls of palisade-shaped sclerenchyma strongly undulate toward the inside and toward the outside, resulting in stellate-undulate facets and a jigsaw puzzle-like pattern on the seed surface except over the raphe and around the micropyle where the cell walls are straight and facets polygonal. Tegmen thin. Embryo tiny, with two rudimentary cotyledons.


In recognition of Léon Paul Choffat (*1849, †1919) for his geological studies of Cretaceous deposits in Portugal.


Cretaceous, Lower Cretaceous
late Aptian – early Albian or older
Type horizon: below the Figueira da Foz Formation


Famalicão (39°42′16″N; 8°46′12″W)

Plant fossil remain

macro- and meso-fossils-embryophytes except wood


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