Juglandicarya simplicarpa Manchester

Plant Fossil Names Registry Number: PFN003661

Act LSID: urn:lsid:plantfossilnames.org:act:3661

Rank: species

Reference for this name: Manchester, S. R. (1987): The fossil history of the Juglandaceae. – Monographs in Systematic Botany from the Missouri Botanical Garden 21: 1–137.

Page of description: 14

Illustrations or figures: fig. 4A–D

Other combinations

Zodicarya simplicarpa (Manchester) M.Deng et Manchester in Deng, Collinson, Zhang & Manchester 2026

Types

Holotype 15740e-4860a, Paleobotany Collections, Florida Museum of Natural History, Gainesville, Florida, USA
Figures: fig. 4A

Note: Holotype inv. no. according to Deng et al. (2026), in Manchester (1987) as "IU 4860".
Holotype refigured in Deng et. al. (2026: pl. 2, figs 1–6).
Paratypes. USNM 313747, 312920 (pl. 2, fig. 12 in Deng et al. 2026), 312944 (pl. 2, fig. 13 in Deng et al. 2026) and 387384 according to Deng et al. (2026: 62). Manchester (1987: 15) listed as paratypes only USNM 313747 (fig. 4B), USNM 312920 (fig. 4C), and USNM 387384 (fig. 4D).

Original diagnosis/description

Locule casts 12–15 mm wide, 13–17 mm high; imprint of surrounding nutshell indicating an outer diam. 14–20 mm wide and 22–40 mm high; wall thickness 1.5–2 mm; two primary lobes of seed well developed, entire or slightly emarginate at base indicating absence or weak development of secióndary septum; shoulder region straight or sloghtly convex; contours of locule cast smotth; nutshell mold rounded and smooth, suggesting little or no ribbing.

Emended diagnosis

[Deng et al. 2026: Fossil Imprint, 82(1): 62–63]
Fruit a unilocular, single-seeded nut. Nut globose-ellipsoidal, 12–22 mm high, 11–14 mm wide; exterior nutshell smooth; primary septum prominent 1.1–1.7 mm thick, at right angles to the plane of separation of the two halves of the nut. Secondary septum absent to weakly developed. Locule bilobed at lower 2/3, entire or slightly emarginate at the base, sometimes giving 4 lobes at the extreme base; apical shoulder region straight or slightly convex; contours of locule smooth without grooves. Anatomy not preserved. Nutshell apparently without lacunae.

Stratigraphy

Paleogene, Paleocene
Fort Union Formation, Tiffanian, middle Paleocene [according to Deng et al. 2026: Fossil Imprint, 82(1): 63]

Locality

United States
Hells Half Acre (UF loc. 15740e), Natrona County, Wyoming [according to Deng et al. 2026: Fossil Imprint, 82(1): 63]

Plant fossil remain

macro- and meso-fossils-embryophytes except wood - angiosperm

Comments

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