Scolecopteris libera Dan D.Li, Jun Wang bis, S.Wan, Pšenička, W.M.Zhou, Bek et Frojdová

Plant Fossil Names Registry Number: PFN000855


Authors: D. D. Li, J. Wang, S. Wan, J. Pšenička, W. M. Zhou, J. Bek & J. Frojdová

Rank: species

Reference: Li, D. D., Wang, J., Wan, S., Pšenička, J., Zhou, W. M., Bek, J. & Votočková-Frojdová, J. (2019): A marattialean fern, Scolecopteris libera n. sp., from the Asselian (Permian)of Inner Mongolia, China. – Palaeoworld 28: 487–507.

Page of description: 488

Illustrations or figures: figs 2–13


Holotype PB22938, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing, China
Figures: fig. 2

Note: Paratypes: PB22939, PB22940, PB22944 (all housed in the Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences).

Original diagnosis/description

Fronds tripinnate. Ultimate fertile pinnae with 10–13 pairs of reproductive pinnules. Pinnules 3.2–4.2 mm long, 2.0 mm wide. Each pinnules with 8–12 synangia, arranged in two rows along the midveins, attached to the middle of the lateral veins with a pedicel. Synangia ca. 1 mm in diameter, 0.7–0.9 mm long, composed of 7–10 sporangia. Sporangia attached to a common pedicel but otherwise free from eachother, exannulate, fusiform with long tips. Trilete microspores circular to oval, 10–20 μm in diameter. Densely microgranulate to microverrucate sculpture on spore exine. Sterile pinnulesof the pecopterid-type, falciform with distinct veins, midveins decurrent, lateral veins bifurcate twice or thrice. Rachises of the Stewartiopteris-type.


The specific name “libera” means free, referring to the fact that the sporangia of a synangium are mostly free from each other, rather than fused laterally as many other species of this genus.


Permian, Cisuralian, Asselian
The uppermost Taiyuan Formation, lowermost Asselian


39°32′25″N, 106°38′03″E, Wuda Coalfield, Inner Mongolia

Plant fossil remain

macro- and meso-fossils-embryophytes except wood


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